Douglas J. Futuyma, State University of New York at Stony Brook, and Mark Kirkpatrick, University of Texas at Austin
Douglas J. Futuyma, State University of New York at Stony Brook, and Mark Kirkpatrick, University of Texas at Austin


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If you are based in the U.S. or Canada and your adoption decision date is looming, please contact Susan McGlew ([email protected]) to request ebook access.

See our brochure for Evolution, Fourth Edition, highlighting the book's special features.

Extensively rewritten and reorganized, this new edition of Evolution features a new coauthor: Mark Kirkpatrick (The University of Texas at Austin) offers additional expertise in evolutionary genetics and genomics, the fastest-developing area of evolutionary biology. Directed toward an undergraduate audience, the text emphasizes the interplay between theory and empirical tests of hypotheses, thus acquainting students with the process of science. It addresses major themes—including the history of evolution, evolutionary processes, adaptation, and evolution as an explanatory framework—at levels of biological organization ranging from genomes to ecological communities.

New in This Edition

  • Genomic perspectives on evolution are strengthened throughout.
  • The content has a stronger focus on human evolution: an entirely new chapter on the topic (Chapter 21, The Evolutionary Story of Homo sapiens), and new examples throughout the book.
  • Many chapters have been rewritten from the ground up.
  • The book has been entirely reillustrated in a clean, contemporary style that enhances the content.
  • A new Appendix, A Statistics Primer, introduces the concept of a probability distribution, reviews how statistics are used to describe populations, looks at how we estimate quantities, and discusses how hypotheses are tested. It ends with a brief overview of two major frameworks of statistical analysis: likelihood and Bayesian inference. Math is kept to a minimum.
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Douglas J. Futuyma is Distinguished Professor in the Department of Ecology and Evolution at the State University of New York at Stony Brook. He received his B.S. from Cornell University and his Ph. D. in Zoology at the University of Michigan with Lawrence Slobodkin. Dr. Futuyma is the author of three previous editions of Evolution, as well as three editions of its predecessor, Evolutionary Biology. He received the 1997 Sewall Wright Award of the American Society of Naturalists and the 2012 Joseph Leidy Award of the Academy of Natural Sciences of Drexel University (Philadelphia). Dr. Futuyma has served as President of the Society for the Study of Evolution, the American Society of Naturalists, and the American Institute of Biological Sciences, and was elected a Fellow of the American Academy of Arts and Sciences in 1996 and the National Academy of Sciences in 2006. He has served as Editor of Evolution and is currently Editor of the Annual Review of Ecology, Evolution, and Systematics. In 2013, he was recognized as Honorary Doctor by the National University of Mongolia. An avid naturalist, his major research interests include evolution of interactions among insects and plants, speciation, and evolution of community structure.

Mark Kirkpatrick is the Painter Centennial Professor of Genetics in the Department of Integrative Biology at the University of Texas at Austin. He received his B.A. in Biology from Harvard in 1978 and his Ph.D. in Zoology from the University of Washington with Monty Slatkin in 1983. Dr. Kirkpatrick has received a Guggenheim Fellowship (1997) and a Poste Rouge Fellowship (France, 1997). He is a Fellow of the American Academy of Arts and Sciences (2008), and of the American Association for the Advancement of Science (2016). Dr. Kirkpatrick received the Sewall Wright Award from the American Society of Naturalists (2014). He has served as Associate Editor of The American Naturalist, Theoretical Population Biology, and Genetics, and on the Editorial Boards of The Annual Review of Ecology, Evolution, and Systematics and Proceedings of the Royal Society of London B. Dr. Kirkpatrick’s research interests are in evolutionary genetics. He has worked on sexual selection, quantitative genetics, speciation, and species ranges. Current research topics include the evolution of sex determination and chromosome rearrangements.

For the Student

Companion Website (evolution4e.sinauer.com)

The Evolution, Fourth Edition Companion Website features review and study tools to help students master the key concepts presented in the textbook. Access to the site is free of charge, and requires no access code. (Instructor registration is required in order for students to access the quizzes.) The site includes the following resources:

  • Chapter Outlines and Summaries: Concise overviews of the important topics covered in each chapter.
  • Data Analysis Exercises: These inquiry-based exercises involve students in working with data and analyzing methods and conclusions from published papers.
  • Simulation Exercises: Interactive modules that allow students to explore many of the dynamic processes of evolution, and answer questions based on the results they observe.
  • Online Quizzes: Quizzes that cover all the major concepts introduced in each chapter. These quizzes are assignable by the instructor.
  • Flashcards: Easy-to-use flashcard activities that help students learn and review all the key terminology introduced in each chapter.
  • The complete Glossary

For the Instructor

(All instructor resources are available to adopting instructors online. Instructor registration is required.)

Instructor’s Resource Library

The Evolution, Fourth Edition Instructor’s Resource Library includes a variety of resources to help you develop your course and deliver your lectures. The IRL includes the following:

  • Textbook Figures and Tables: All the figures (including photographs) and tables from the textbook are provided as JPEGs, reformatted and relabeled for optimal readability when projected.
  • PowerPoint Presentations: For each chapter, all of the figures and tables are provided in a ready-to-use PowerPoint presentation that includes titles and full captions.
  • Answers to the textbook end-of-chapter Problems and Discussion Topics
  • Quiz Questions from the Companion Website
  • Data Analysis and Simulation Exercises from the Companion Website, with answers

Online Quizzing

A set of online quizzes is available via the Companion Website. These quizzes can be assigned or released for student self-study, at the instructor’s discretion. Instructors can also add their own questions to the quizzing system, to create custom quizzes. Results can be viewed online or downloaded for use in gradebook programs. (Instructor registration is required for student access to the quizzes.)

If you have adopted this text for course use (within the U.S. or Canada) and are interested in the instructor’s supplements that accompany the text, please contact [email protected]. Outside the U.S. or Canada? Check the orders and returns page for the distributor in your region.