Richard C. Brusca, University of Arizona, and Emeritus, Arizona–Sonora Desert Museum, Wendy Moore,...

Richard C. Brusca, University of Arizona, and Emeritus, Arizona–Sonora Desert Museum, Wendy Moore, University of Arizona, and Stephen M. Shuster, Northern Arizona University, with 22 contributors

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See our brochure for Invertebrates, Third Edition here.

In the 12 years since publication of Invertebrates, Second Edition, fundamental shifts have occurred in our understanding of the origins and evolutionary relationships among protists and animals. These changes are largely due to the explosion of molecular...

See our brochure for Invertebrates, Third Edition here.

In the 12 years since publication of Invertebrates, Second Edition, fundamental shifts have occurred in our understanding of the origins and evolutionary relationships among protists and animals. These changes are largely due to the explosion of molecular phylogenetics and evo-devo research, emergence of the new field of animal genomics, major fossil discoveries in China, Australia, and elsewhere, and important new embryological and ultrastructural studies. As a result:

  • New phyla have been described (e.g., Micrognathozoa, Xenacoelomorpha).
  • Old phyla have been collapsed into others (e.g., Sipuncula and Echiura are now placed within Annelida; acanthocephalans are now known to be highly modified, parasitic rotifers).
  • Phyla once thought to be deuterostomes are now part of the protostome clade (e.g., Chaetognatha, Phoronida, Bryozoa, Brachiopoda).
  • The Protostomia has been reorganized into two major clades known as Ecdysozoa and Spiralia.

For each of the 32 currently recognized phyla, Invertebrates, Third Edition presents detailed classifications, revised taxonomic synopses, updated information on general biology and anatomy, and current phylogenetic hypotheses, organized with boxes and tables, and illustrated with abundant line drawings and new color photos. The chapters are organized around the “new animal phylogeny,” while introductory chapters provide basic background information on the general biology of invertebrates. Two new coauthors have been added to the writing team, and 22 additional invertebrate zoologists have contributed to chapter revisions. This benchmark volume on our modern views of invertebrate biology should be in every zoologist’s library.

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Richard C. Brusca is Executive Director, Emeritus of the Arizona-Sonora Desert Museum and a Research Scientist at the University of Arizona. Rick is the author of nearly 200 research publications and 13 books, including the popular field guides Common Intertidal Invertebrates of the Gulf of California, A Seashore Guide to the Northern Gulf of California, and A Natural History the Santa Catalina Mountains, with an Introduction to the Madrean Sky Islands. He has been the recipient of more than 100 research grants from the National Science Foundation, NOAA, the National Geographic Society, the David and Lucile Packard Foundation, and many other agencies and foundations. He has served on panels for the National Science Board, National Science Foundation, Smithsonian Institution, PEW Program in Conservation and the Environment, Public Broadcasting Service, IUCN Species Survival Commission, and many others. Rick has also served on many environmental non-profit boards, in the U.S. and abroad, and he has organized and conducted field expeditions throughout the world, on every continent.  He is an elected Fellow in the American Association for the Advancement of Science (AAAS), the Linnean Society of London (FLS), and the California Academy of Sciences.

Wendy Moore is Assistant Professor in the Department of Entomology at the University of Arizona and Curator of the University of Arizona Insect Collection. Her degrees were earned at Vanderbilt University (B.S., General Biology), the College of Charleston (M.S., Marine Biology), and the University of Arizona (Ph.D., Entomology/Ecology and Evolutionary Biology). Dr. Moore’s long-term research interest is the evolution of biotic diversity—especially the evolution of symbiotic lifestyles and how major biotic, climatic, and tectonic events may have influenced the timing and patterns of diversification. Much of her current research is on the carabid beetle subfamily Paussinae, many species of which are obligate symbionts with ants. She is also deeply committed to collections care and enhancement, and the use of bioinformatics to make collections-based data widely available to diverse user communities.

Stephen M. Shuster is Professor of Invertebrate Zoology and Curator of Marine Invertebrates and Molluscs at Northern Arizona University. He earned a B.S. in Zoology from the University of Michigan, an M.S. in Biology from the University of New Mexico, and a Ph.D. in Zoology from the University of California, Berkeley. The author or coauthor of over 75 journal articles, encyclopedia entries, book reviews, and contributed book chapters, Dr. Shuster collaborated with Michael J. Wade on the book Mating Systems and Strategies (Princeton University Press, 2003). His research broadly concerns mating system evolution, male and female reproductive behavior, community and ecosystem genetics, and the population biology of marine organisms. His recent work focuses on the measurement of selection within and among species, and the maintenance of genetic variation in natural populations of marine crustaceans and terrestrial arthropods.

Contributing authors include:
Jesús Benito · Sarah Cohen · Gonzalo Giribet · Rick Hochberg · Gustavo Hormiga · Reinhardt Møbjerg Kristensen · David Lindberg · Carsten Lüter · Joel W. Martin · Alessandro Minelli · Rich Mooi · Ricardo Cardoso Neves · Claus Nielsen · Fernando Pardos · Winston Ponder · Greg Rouse · Scott Santagata · Andreas Schmidt-Rhaesa · George Shinn · Martin Vinther Sørensen · S. Patricia Stock · Katrine Worsaae

“Vives félicitations aux auteurs et éditeurs de cet excellent et brillant volume.”
—Pierre Jolivet, L'Entomologiste

“A wonderful book, and one remains amazed at the perfection of this edition, full color, and the precision and detail of the figures. … A masterpiece that will dominate the phylogeny for many, many years.”
—Pierre Jolivet, L'Entomologiste, Paris (from the original French)

“Through all my years of full-time teaching (starting in 1992) I consistently had a copy of the most recent text by Brusca on my shelf as a reference book, but this edition, IMO, finally nailed it as a teaching text. … This text, now co-authored by Richard Brusca, Wendy Moore, and Stephen Shuster, is much more readable and student friendly than past editions. In fact, it's downright fantastic. … The overall approach is comparative anatomy. … Before I decided whether to adopt this textbook I took a careful look at its treatment of several taxa that have been problematic in the past. … In every case this book's treatment of these and all taxa I have reviewed includes up-to-date systematics. I am also pleased to see the updated treatment and taxonomy of all eukarya in Chapter 3. … I am impressed by the layout of each chapter. In most invert textbooks taxonomic information is relegated to the back of a chapter where students may or may not give it only a cursory glance. Brusca's team, however, puts the taxonomy right up front and [provides] a brief history of the taxonomy of each major group. This provides an interesting context that allows students to see how science works as we uncover new observations and apply new methods, in this case improving the taxonomy and systematics of animal life. The chapter layout should also make this an easy text to teach from and learn from. … The addition of color photos to supplement the excellent stipple line drawings is also a hallmark of this title. I can't wait to get started using this textbook in the classroom. … 5 solid stars for what I believe is the gold standard of textbooks in this field of study.”

“I am so thrilled that we finally getting a third edition of Invertebrates! I’ve been teaching invertebrate zoology for over 15 years and this text is superior to any other on the market!”
—Tamara J. Cook, Sam Houston State University

For the Instructor (Available to Qualified Adopters)

Instructor’s Resource Library
Available to qualified adopters, the Instructor’s Resource Library for Invertebrates, Third Edition contains an extensive collection of images for use in teaching the course:

  • Textbook Figures and Tables: All of the textbook’s figures and tables are included as both high- and low-resolution JPEGs, for easy use in presentation software, learning management systems, and assessments. New for the Third Edition, this now includes all of the textbook's photographs.
  • Supplemental Photo Collection: This collection of over 900 photographs depicts organisms that span the entire range of phyla covered in the textbook.
  • PowerPoint Presentations: Two ready-to-use PowerPoint presentations are provided for each chapter of the textbook: one that contains all of the textbook figures and tables, and one that contains all of the relevant photos from the supplemental photo collection.

If you have adopted this text for course use (within the U.S. or Canada) and are interested in the Instructor's Resource CD, please contact [email protected]. Outside of the U.S. or Canada? Check the orders and returns page for the distributor in your region.

Errata for the First Printing

Preface to the Third Edition, p. XIV, left column, 8 lines from the bottom
Text correction: The basal, diploblastic phyla have been shuffled about near the base of the Metazoa tree, and they might continue to shuffle about for a bit longer.

Preface to the Third Edition, p. XIV
Text correction:
R.C.B., W.M., S.M.S.
Tucson, Arizona
December 2015

Chapter 1, p. 9, right column, last sentence of first paragraph
Text correction: About 15,000 to 20,000 new species are named and described by biologists each year, most of them invertebrates (mainly insects).

Chapter 1, p. 24, left column, line 22 from the top
Text correction: Imagine what will happen to the giant fig trees of the tropics (Ficus is the most widespread plant genus in the tropics) as they lose the single parasitic wasp varieties that pollinate each of the 900 or so species.

Chapter 3, p. 114
Text correction, Figure 3.42 title: Phylum Parabasalida.

Chapter 5, p. 199
The heading “Settling and Metamorphosis” should be a subheading of “Indirect Development” on p. 197.

Chapter 7, p. 265, fifth line from the top
Text correction: There are about 13,200 living described species of cnidarians.

Chapter 7, p. 283
Text correction, Figure 7.10, end of caption: (C) Vellela skeletons in beach wrack, coast of Washington State (USA).

Chapter 13, p. 517
Text correction, Figure 13.48 caption: Stacks of Crepidula fornicata, a slipper shell (Caenogastropoda) displaying sequential hermaphroditism.

Chapter 20, p. 71
Table 20.1: The first column heading should read “Phylum/Subphylum” (not just “Phylum”).

Chapter 25, p. 973, left column, first order
Text correction: ORDER COMATULIDA. Feather stars and their allies. Stem usually lost following postlarval stage; feather stars retain only the most proximal stem element bearing anchoring hooklike cirri; species retaining the stalk as adults lack cirri. (e.g., Antedon, Bourgeticrinus, Comatula, Mariametra, Tropiometra)

Running heads for Chapters 23 and 24 should read “ARTHROPODA.”